Professor Bryan Campbell Clarke FRS, was a British geneticist, latterly professor emeritus of genetics at the University of Nottingham. Clarke is particularly noted for his work on apostatic selection and other forms of frequencydependent selection, and work on polymorphism in snails, much of it done during the 1960s. Later, he studied molecular evolution. He made the case for natural selection as an important factor in the maintenance of molecular variation, and in driving evolutionary changes in molecules through time. In doing so, he questioned the overriding importance of random genetic drift advocated by King, Jukes, and Kimura. With Professor James J Murray Jnr , he carried out an extensive series of studies on speciation in land snails of the genus Partula inhabiting the volcanic islands of the Eastern Pacific. These studies helped illuminate the genetic changes that take place during the origin of species.
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Source: Wikipedia